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quill (kwĭl)n.1. The hollow stemlike main shaft of a feather. Also called calamus.2. Any of the larger wing or tail feathers of a bird.3. A writing pen made from the shaft of a feather.4. Music a. A plectrum for a stringed instrument of the clavichord type.b. A pipe having a hollow stem.5. A toothpick made from the stem of a feather.6. One of the sharp hollow spines of a porcupine or hedgehog.7. A spindle or bobbin around which yarn is wound in weaving.8. A hollow shaft that rotates on a solid shaft when gears are engaged.v. quilled, quill·ing, quills v.tr.1. To wind (thread or yarn) onto a quill.2. To make or press small ridges in (fabric).v.intr. To practice the art or craft of quilling.[Middle English quil.]